Placentation

Placentation
Placentation resulting from cleavage at various gestational ages
Latin placentatio

In biology, placentation refers to the formation, type and structure, or arrangement of placentas. The function of placentation is to transfer nutrients from maternal tissue to a growing embryo. Placentation is best known in pregnant female mammals (eutheria), but also occurs in other animals, eggs (yolk sac placentation) and flowering plants.

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Placentation in mammals

In placental mammals, the placenta forms after the embryo implants into the wall of the uterus. The developing fetus is connected to it via an umbilical cord. Animal placentas are classified based on the number of tissues separating the maternal from the fetal blood. The placentation types found in animals are:

In hemochorial placentation maternal blood comes in direct contact with the fetal chorion, which it does not in the other two types.[2] It may avail for more efficient transfer of nutrients etc., but is also more challenging for the systems of gestational immune tolerance to avoid rejection of the fetus.[3]

During pregnancy, placentation is the formation and growth of the placenta inside the uterus. It occurs after the implantation of the embryo into the uterine wall and involves the remodeling of blood vessels in order to supply the needed amount of blood. In humans, placentation takes place 7–8 days after fertilization.

In the case of twins, dichorionic placentation refers to the presence of two placentas (in all dizygotic and some monozygotic twins). Monochorionic placentation occurs when monozygotic twins develop with only one placenta and bears a higher risk of complications during pregnancy. Abnormal placentation can lead to an early termination of pregnancy, for example in pre-eclampsia.

Placentation in plants

In flowering plants, placentation occurs where the ovules are attached inside the ovary.[4] The ovules inside a flower's ovary (which later become the seeds inside a fruit) are attached via funiculi, the plant part equivalent to an umbilical cord. The part of the ovary where the funiculus attaches is referred to as the placenta.

In botany, the term placentation most commonly refers to the arrangement of placentas inside a flower or fruit. Plant placentation types include:

See also

References

  1. ^ Claim for guinea pigs, rabbits, mice, and rats taken from: Thornburg KL, Faber JJ (October 1976). "The steady state concentration gradients of an electron-dense marker (ferritin in the three-layered hemochorial placenta of the rabbit". J. Clin. Invest. 58 (4): 912–25. doi:10.1172/JCI108544. PMC 333254. PMID 965495. http://www.pubmedcentral.nih.gov/articlerender.fcgi?tool=pmcentrez&artid=333254. 
  2. ^ thefreedictionary.com > hemochorial placenta Citing: Dorland's Medical Dictionary for Health Consumers. Copyright 2007 by Saunders
  3. ^ Elliot, M.; Crespi, B. (2006). "Placental invasiveness mediates the evolution of hybrid inviability in mammals". The American naturalist 168 (1): 114–120. doi:10.1086/505162. PMID 16874618.  edit
  4. ^ "Flowers" At: Botany Online At: University of Hamburg Department of Biology. (see External links below).

External links